Polymorphism
The hindwings are bright red across most of the range of the species, but in the northwest of the range the species is polymorphic, with red, orange, or yellow hindwings. The colour is part of the species’ warning coloration (aposematism), signalling to possible predators that the species is distasteful, in its case protected by harmful chemicals. It is unusual for such warning signals to vary, as a single pattern is most likely to be an effective deterrent.
The polymorphism is created by two genes. Each gene has two alternative forms (alleles), one dominant to the other. One of the genes is epistatic to the other. One gene gives red if the dominant allele R is present, else orange (if homozygous for the recessive allele r); the other gene gives yellow if homozygous for its recessive allele (yy), regardless of the alleles of the red gene. Thus the Y allele is required for red or orange to appear. The Y allele may enable metabolism of the yellow pigment to orange, which the R allele can then process further to give red, which would explain the epistasis.