March Flies
family of suborder “Nematoceran Flies“
1 family, 2 species
Bibionidae, or March flies, are medium-sized flies known for their black, brown, or rusty bodies and thick legs. Their larvae are herbivores and scavengers, feeding on dead vegetation or plant roots. Some species can be found in compost. Adult Bibionids are short-lived and do not eat, relying on food from their larval stage. Fossil records indicate that Bibionids have been around since at least the Jurassic period. They are important pollinators and their larvae contribute to the accumulation of humus in soil. Some larvae can be pests to agricultural crops.
Hierarchy
species of family “March Flies“
1 species
Description#
Bibionidae are medium-sized flies with a body length from 4.0 to 10.0 mm. The body is black, brown, or rusty, and thickset, with thick legs. The antennae are moniliform. The front tibiae bear large strong spurs or a circlet of spines. The tarsi are five-segmented and bear tarsal claws, pulvilli, and a well developed empodium. The wings have two basal cells (posterior basal wing cell and basal wing cell), but are without a discoidal wing cell. R4+5 is simple or branched; at most, only three branches of R developed. The leading edge wing veins are stronger than the weak veins of the trailing edge.
Biology#
Bibionid larvae grow up in grassy areas and are herbivores and scavengers feeding on dead vegetation or living plant roots. Some species are found in compost. Larvae are sometimes found in pockets in which sometimes up to 200 specimens have been counted. Adults of most Plecia and some species of Bibio do not eat, but subsist solely on the food taken in during the larval stage and drop steadily when in flight until they are a few inches above the ground, hovering slowly. Adult-stage bibionids are quite short-lived, and some species of Plecia (lovebugs) spend much of their adult lives copulating. The male and female (lovebugs) attach themselves at the rear of the abdomen and remain that way at all times, even in flight. Adults swarm after synchronous emergence, sometimes in enormous numbers.
Fossil record#
Bibionids have the most extensive fossil record of any dipteran family. Fossil bibionids are known questionably from the Jurassic, while some forms from the early part of the Upper Cretaceous look quite similar to modern species. Bibionid flies are very abundant among insect fossils from the Tertiary period, and many species have been described, although often based on highly fragmentary material. Most fossil species are easily identified with extant genera. In particular, the genera Plecia and Bibio are abundant among Tertiary fossils. Fossils from Europe include many specimens of the mainly tropical genus Plecia which is today entirely absent from Europe, demonstrating a warmer climate during the Tertiary.
Genera#
Bibio Geoffroy, 1762 Bibiodes Coquillett, 1904 Bibionellus Edwards, 1935 †Cascoplecia Poinar, 2010 †Clothonopsis Hong & Wang, 1987(dubious taxon) Dilophus Meigen, 1803 Enicoscolus Hardy, 1968 †Fushunoplecia Hong, 2002 †Lithosomyia Carpenter, 1986 †Mesopleciella Rohdendorf, 1946 Penthetria Meigen, 1803 Plecia Wiedemann, 1828
Economic importance#
For those species where the adults feed, they do so on the nectar of flowers of fruit trees and especially on flowers of umbelliferous plants, often swarming in mass flights in spring. Adults are important pollinators. Larvae play an important role in formation and accumulation of humus in soil. Some larvae are serious plant pests, especially of pasture land and other agronomic crops including cereal crops, vegetables, forage crops, and seedlings of many other plants.
Images#
Diptera.info BugGuide
External links#
multiple images including photomicrographs at Encyclopaedia of Life Plecia nearctica, lovebug on the UF / IFAS Featured Creatures Web site
Bibionidae, or March flies, are medium-sized flies known for their black, brown, or rusty bodies and thick legs. Their larvae are herbivores and scavengers, feeding on dead vegetation or plant roots. Some species can be found in compost. Adult Bibionids are short-lived and do not eat, relying on food from their larval stage. Fossil records indicate that Bibionids have been around since at least the Jurassic period. They are important pollinators and their larvae contribute to the accumulation of humus in soil. Some larvae can be pests to agricultural crops.
Ancestry Graph
Further Information
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